MEMOIRS OF THE NATIONAL ACADEMY OF SCIENCES Volume XII FIRST MEMOIR WASHINGTON 1914 '[ NATIONAL ACADEMY OF SCIENCES. Volume XII. FIRST MEMOIR. MONOGRAPH OF THE BOMBYCINE MOTHS OF NORTH AMERICA, INCLUDING THEIR TRANSFORMATIONS AND ORIGIN OF THE LARVAL MARKINGS AND ARMATURE. PART III. FAMILIES CERATOCAMPID^ (EXCLUSIVE OF CERATOCAMPIN^), SATURN I IMS, HEMILEUCID^), AND BRAHMiEID^E. BT ALPHEUS SPRING PACKARD, EDITED BY THEODORE D. A. COCKERELL, BEING A CONTINUATION OF PREVIOUS INVESTIGATIONS PUBLISHED AS VOLUME VII (FIRST MEMOIR) AND VOLUME IX (SECOND MEMOIR) OF THE MEMOIRS OF THE NATIONAL ACADEMY" OF SCIENCES. TABLE OF CONTENTS. 1. Preface vn 2. Introduction IX 3. Family Ceratoeampidee 1 4. Family Hemileucidse 77 5. Family Saturniidse 151 6. Hybrid Saturniidfe 265 7. Parasites 267 8. Genera of uncertain family position 269 9. Brahmseidse 271 v PREFACE. The manuscript and notes for this final volume of my husband's (Alpheus Spring Packard) Monograph of the Bombycine Moths are printed, with the exception of editorial additions, exactly as he left them at the time of his death on February 14, 1905. He had been working on them during the leisure intervals of college duties, and it was one of the last wishes he expressed that the National Academy of Sciences might consent to print the unfinished part, since they had already printed Parts I and II. My husband was fully aware of the incomplete condition of this later part, and had expected to spend much time in finishing it; he had also hoped to revisit the British Museum in order to work from the types in the collections there. My husband's scientific friends who have been consulted in regard to printing have agreed that although these parts are incomplete, yet the valuable results of so many years of labor should be put into accessible and permanent form. The accomplishment of this purpose is due above all to the labors of Prof. Theodore D. A. Cockerell, of the University of Colorado, who has most generously given Ids time to editing this volume. Without his kind and able assistance there would have been further delay of publi- cation. I must leave to Prof. Cockerell all acknowledgments to those who have kindly helped him by supplying illustrations, etc. Our obligations are great to both Dr. Samuel Henshaw, of the Museum of Comparative Zoology, and to Miss Caroline G. Soule, of Brookline. We must ask my husband's many friends who furnished him so generously either with specimens or the results of their own observations to realize that through ignorance we can not make the proper acknowledgments, as he would have been careful to do. Elizabeth Walcott Packard. Andover, Mass., July, 1912. INTRODUCTION The manuscripts left by Dr. Packard show that he contemplated what would have amounted to a monograph of the Saturnioid moths of the world. Beginning with the North American forms, he soon found it necessary to make comparisons with those of other regions, and during his last years was actively engaged in examining the many exotic genera. As all lepidopterists know, he was led to novel and interesting conclusions regarding the classification of these insects, parts of which were published in short papers before his death. It was in the mono- graph, however, that he hoped to fully expound and justify his opinions, while at the same time making known the life histories of the Saturnioids in a manner never before attempted. As will be seen from the following pages, great progress had been made, but very much remained to be done. The latest writings of Dr. Packard show that he was actively seeking new light and continually acquiring new information which led him to modify the details of his taxo- nomic scheme. It is impossible to say how much the work now presented would have been remodeled by him had he lived to this day, but it is at least certain that it would have under- gone much modification and especially amplification. The editor has left the work exactly as it came from Dr. Packard's hands except in the following particulars: (1) The arrangement is that of the editor, following, however, the order of genera preferred by Dr. Packard, so far as could be ascertained from a study of his writings. (2) A few obvious slips of the pen have been corrected, and some very incomplete manu- scripts have been omitted. (3) Additions have been made, but in every case indicated by square brackets. Such additions are the work of the editor unless some other signature is appended. It was evidently impossible to complete the work on the lines laid down by Dr. Packard, but it seemed practicable and desirable to bring the account of the North American species up to date. In the case of the exotic genera only enough has been added to bring out inter- esting points or give a general idea of the extent and distribution of the groups. A few genera have been entirely omitted, either because Dr. Packard made no reference to them or because they did not appear to belong to the families included in the work. It has often been necessary to add bibliographical references, and when these could not be verified from the original works Kirby's Catalogue was the source of information. The editor is greatly indebted to many kind friends and correspondents who did every- thing in their power to aid in the work. Dr. H. G. Dyar supplied much valuable material and answered many questions, and also selected from the collection in the United States National Museum a splendid series of moths to be photographed, enabling us to illustrate for the first time and from the original types many American species. Mr. J. H. Watson, of Manchester, England, aided by the loan of literature, by answering questions, and especially by supplying a very fine series of photographs representing numerous genera discussed by Dr. Packard, but represented only by drawings of venation, or not at all, among the illustrations left by him. Dr. J. McDunnough, of Decatur, 111., has kindly prepared descriptions of several North Ameri- can species and has sent photographs of rare forms in the Barnes collection. Dr. L. O. Howard and his associates in the Bureau of Entomology, United States Department of Agriculture, have placed at my disposal the numerous manuscript records of the bureau. Mr. Jacob Doll, of the Brooklyn Museum, very kindly sent photographs of the North American species in his charge not otherwise represented on our plates. Dr. Glover M. Allen copied several descrip- tions inaccessible to me and looked up a number of doubtful references. Other assistance is mentioned at various places in the text, but it has been impossible to acknowledge in any sufficient way the results of Mrs. Packard's always active interest and cooperation. On the financial side we are greatly indebted to the trustees of the Bache fund for a grant of $200 toward the cost of the preparation of the plates. T. D. A. COCKERELL. University of Colorado, Boulder, Colo., August, 1912. IX THE BOMBYCINE MOTHS OF NORTH AMERICA, PART III. By Alpheus Spring Packard. Superfamily SYSSPHINGINA Packard. Family CERATOCAMPID^ (Harris) Packard.1 Subfamily 1. Ceratocampin.e Grote. (See Part II, Memoirs of the National Academy of Sciences, vol. 9.) Subfamily 2. Agliin^e Packard. Agliinx Packard, Proc. Amer. Phil. Soc. [XXI (1893), p. 139]. Agliinse Packard, Ann. and Mag. Nat. Hist. [(6), XI (1893), pp. 172-175]. Head rather large, unusually narrow between the eyes; the vestiture rather long, but not shaggy or radiating. Antennae of 6" varying greatly, either bipectinated, with short very densely ciliated branches (Arsenura), or the branches still shorter, forming almost simple teeth, much ciliated (Dysdse- monia), or very widely pectinated to the tip (Aglia and Polythysana), or moderately bipectinated nearly to the tip (Bathyphlebia) . In 9 either with very short pectination (Aglia, Polythysana), or entirely simple (Arsenura and Dysdsemonia). Palpi very constant in form, being unusually well developed, large, compressed, ascending, passing beyond the front, distinctly 3-jointed, reminding one often of the Sphingidae. * Maxilla? unusually well developed, though slender and short; in Dysdsemonia the two appendages are united for about half their length, and they are as long as the head is broad, or about one-third as long as the S antenna?. Thorax rather stout, abdomen conical. Wings often large; fore wings with the costa much curved on the outer half; the apex either somewhat produced and rounded, or broad and square (Dysdsemonia); outer edge normal, moderately full (Aglia) or slightly falcate (Polythysana and Arsenura), or much so, and the outer edge deeply excavated and scalloped in Dysdsemonia. Hind wings either normal, with the outer edge either full and rounded (Aglia, Polythysana) ; or with a long angular projection at the end of veins III2 and III3 (Arsenura), or in Dysdsemonia, with a rather long tail widening at the square, flaring end, supported by veins III2 and III3. Venation: Fore wings, with 11 veins; vein III2 independent, entirely detached from its original vein or stalk (Arsenura and Dysdsemonia) ; in Aglia and Polythysana the vein is less detached. In all the genera examined the vein IL. arises within the origin of the anterior discal vein, while the origin of this vein, in Aglia, is situated nearer to the discal vein than in any other genus of the group. Another almost diagnostic character of the subfamily is the small size of the discal cell; in all the genera the outer end or side is situated well inside of the middle of the wing; it is widest in Dysdxmonia, narrowest in Arsenura. Hind wings with eight veins; the discal cell is small and short, except in Polythysana, where it is two-thirds as long as the entire wing along the median vein (IV), the genus being very aberrant in this respect. The common line formed by the two discal veins taken together is very oblique (Arsenura) ; in Aglia much less so. In the venation of the hind wings Aglia is in all important respects much like Arsenura and Dysdsemonia. Legs rather large, long, and slender. ' [According to Art. 5 of the International Rules of Zoological Nomenclature, this family must be called Citheroniida? Dyar, 1894, and the sub- family must be CitheronUnse.] 83570°— 14 1 1 2 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, Larva in its last stage spineless, smooth; in the early stages with 2 to 6 thoracic spines, and a median double spine on eighth abdominal segment. Pupa like that of Eacles [Packard, Psyche, Feb. 1902, p. 305]. Judging by the adult characters the Agliinse may be divided into three groups, the first being represented by Arsenura, which appears to be the most generalized form, its ally, Dysdse- monia, being more specialized, having undergone reduction in its maxillae, but a higher degree of specialization as seen in its tailed hind wings, and the diaphanous discal and secondary spots, as also the highly modified, almost simple, but densely ciliated antennas. The second group is represented by Aglia, and apparently Baihyphlebia. The third by Polythysana. It is quite apparent that Aglia is an offshoot of the Arsenura phylum. As to the origin of Polythysana, that remains to be settled after we have a knowledge of the larva and all its stages. Until its transformations became known and comparisons made with those of Sphingidae and Ceratocampidae, the European Aglia tau was allowed to remain in the family Saturniidse, as formerly understood. ARSENURA Duncan. Phalxna-Attacus Cramer, Papillons Exotiques, III, t. 197A, 1782. Arsenura Duncan, Naturalists Library, p. 125 (no description), "1837." Rhescyntis group 2, Walker, Cat. Lep. Het. Br. Mus., VI, p. 1323, 1855. Arsenura Drtjce, Biologia Centrali-amer., Lep. Het., I, 1886. Arsenura Kirby, Syn. Cat. Lep. Het., 1, p. 769, 1892. Imago. — young and full-fed larva.) Burmeister, H., Description phys. Rep. Argentine, Lep., Atlas, Liv. 1 and 2, p. 47, PI. XXI, figs. 1, 1A, 1879-80. Peters, H. T., Die Heteroceren-Raupen, p. 9, Taf. VI, fig. 9, 1898-1901. (Young, half grown, and full fed or last stage.) Imago. — 1 £19. Body and wings uniformly fawn brown, the S a little darker than 9 . Head and palpi much darker than the rest of the body. Fore and hind wings not scalloped on the edges. Fore wings with a broad diffuse basal line situated much nearer the discal streak than the base of the wing; on the outer or hinder half of the wing it is curved outward. Discal spot an indistinct broad line alike on both wings. Extradiscal line dark fawn, oblique, not waved or scalloped, slightly excurved on inner edge of wing, ending on the costa on the outer third of the wing. Halfway between the end of this line and the apex of the wing is a double dark-brown irregular costal streak, from which passes a white line to the apex. A sinuous white line edged more or less distinctly on each side curving outward in a large scallop toward the apex and behind the double blackish spot; it makes two similar scallops in the middle of 6 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, the wing. In the apical region are two linear white scallops, the hinder one bordered with an oblique chestnut-brown streak reaching the extradiscal line. Beyond the extradiscal line the wing is clear fawn brown, while within the extradiscal line the wing (except on the inner edge) in hoary or gray, much less so in $ than in 9 , with the angle in male well marked. Hind wings colored and marked like the fore wings; no basal line. The extradiscal line broad, straight, dark, and touching the submarginal scalloped line on the inner edge of the wing. This line is made up of six scallops, with dark points between them, the first, fourth, and fifth points three times larger than the second ami third; in the S the points are more nearly equal in size and form a more continuous blackish line, with scattered white scales. Beneath slightly paler, but the lines and other markings are as on the upper side though less distinct, and the dark submarginal line is nearly effaced in the middle of the wing away from the costal and inner edge of the wing. Legs: The femora and tibiae dark, like the palpi, but the tarsi are much paler. Abdomen with a row of six lighter oval triangular spots encir- cled with dark on each side. Expanse of fore wings, $ 120 mm.; 9 142 mm. Length of fore wings, S 62 mm.; ? 70 mm. Breadth of fore wings, S 30 mm.; $ 36 mm. Length of hind wings, S 46 mm.; and frauenfeldi (Felder) are aberrant forms of 0. venusta Walker. Larva. — Our knowledge of the larva of this genus is derived from a colored drawing in the British Museum, of which fig. 14, PI. XXXI, is a copy, which I was kindly allowed by Sir George Hampson to have made. It is labelled " 0. frauenfeldi." On comparing this with an alcoholic example of Aglia tau, and the excellent figure by Prof. Poulton, which evidently well represents its characteristic attitude when at rest, the resemblance between the larvae of the two genera in their final stage is striking. In Cercophana the body is smooth and unarmed; the segments apparently not convex, and smooth; the head is small and the two anterior thoracic segments small, while apparently the tergum of the metathoracic segment (unless it be the first abdominal) is prolonged into a long median process, which overhangs the retracted head and two anterior thoracic segments. Also the suranal plate is greatly prolonged into a sharp conspicuous process. Along the sides is a row of six long black setae which extend downward and backward. The body is brownish 28 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, or bronze along the back, green on the side and beneath, while a conspicuous yellow lateral line extends from the tip of the anterior to that of the posterior horn or projection. It has no lateral eye-like red spot on the side of the first abdominal segment. The larva, however, so far as we can judge from a figure alone, is evidently closely alhed to Aglia tau, as is the moth. It will be a matter of the greatest interest to have the larval history of this genus worked out; we should expect that its freshly hatched larva is armed with spines as in Aglia, which are discarded at the last molt. It is a matter of the utmost importance that the entomologists of Chile should secure, the eggs and rear the larva, securing the larva; in all stages and the pupa, and to ascertain whether the pupa is subterranean or protected in a cocoon. CERCOPHANA MIRABILIS Rothschild. Cercophana mirabilis Rothschild, Nov. Zool., II, p. 46, PL X, figs. 6, 7, 1895. Imago. — "The most obvious differences which separate this species at a glance from any of the varieties of E. venusta (Walk.) are its small size (barely half that of venusta), strongly dentated margins to all wings, and absolutely tailless hind wings in both sexes. "Male: Fore wings deep rufous chocolate, a large round white spot situated at apex of cell, beyond which is a transverse bar of darker chocolate. Hind wings orange yellow, with the outer third reddish chocolate, and a central narrow transverse band of the same color. "Underside similar, but all the colors and markings more mixed and indistinct. "Female: Fore wings reddish gray, with a dull yellow round spot at the apex of cell, between which and the base of the wing are two indistinct red transverse fines, and beyond the cell again are two broader and more distinct ones. "Hind wings reddish gray, more brown toward the margins, and crossed by two very indistinct transverse fines. Under side identical. "Expanse, o* 1 i inches = 38 mm.; 9 2 inches = 51 mm. "Hab. Chile (in coll. Staudinger)." (Rothschild.) This is probably the type of quite a distinct genus; the fore wings are not falcate, and the hind wings besides being tailless are quite different in shape. [It is said to look like an Orgyia.] [NEOCERCOPHANA Izquierdo, 1896. Plate CXI, figs. k. 1. The type of this genus is N. philippii Izquierdo, 1896, from Chile. Izquierdo describes the metamorphoses.] Subfamily Bun^in^: Packard. [Bunaeinx Packard, Psyche, February, 1902, p. 306.] Head: When denuded the front is somewhat shield-shaped or subtriangular, moderately wide, a little wider than one eye seen from in front, narrowing a little toward the oral region. Antennas of S bipectinate, either with rather short pectinations on basal three-fourths, leaving the tip filiform (Imbrasia) or pectinated to the tip and more or less plumose, the pectinations being unusually long and slender; those of ? subsimple (or simple). Palpi three-jointed. Wings large and broad; fore wings with 11 veins, broad, large, as a rule about one-half as wide as long; the costal edge more or less curved; the apex either subacute (Lobobunsea) or acute (Bunaea) ; the outer edge entire ( Vsta) or excavated, and in certain genera (Lobobunsea and Bunaea) scalloped. Hind wings either normal, the apex squarish, the outer edge slightly convex (Vsta), or the apex is rounded, and there is a decided angle in the outer edge (Girina) or a short tail (Imbrasia), but in the other genera the outer edge is convex, full, though Gynanisa ethra has a short tail. no. i. BOMBYCINE MOTHS OF NORTH AMERICA— PACKARD. 29 Vein III2 is never detached so as to form an independent vein; there is only a tendency to it in Nudaurelia, the tendency being most marked in [Acanihocampa] belina. The venation is very uniform. The number of veins is invariably eight. Fore wings : The discal cell rather narrow; the two discal veins form a line wliich is situated either in the middle (Usta, Salassa) or a little beyond the middle of the wing (Bunaea), and the line varies in being curved or bent outward or inward. It is situated farthest outward in Bunaea. Vein II, [III, in revised nomenclature] arises before the origin of the stalk of II3 and Uit in the outer third of the discal cell (in Vsta, Cirina, Imbrasia, Antherina, Nudaurelia, Gynanisa, Lobobunsea, and Salassa). The same vein in Thyella, [Acanihocampa] belina and Bunaea arises at the end of the discal cell in front of the anterior discal vein. Hind wings: Vein III2 semiindependent in Imbrasia and Antherina; in [Acanihocampa] belina it becomes almost entirely independent, more so than in any other genus of the group. [The original definition of Bunaeinse, in the place sited, is as follows: "Antennae bipectinate, tip filiform, or pectinated to tip. Vein III2 is never detached so as to form an independent vein. Wings usually very large, and in the more specialized genera closely approaching the Saturnian Anthersea (a case of parallelism or convergence), but the larvae are entirely different, not spinning a dense cocoon and being armed with stout long spines (in certain genera spinulated), instead of soft tubercles crowned with several small short spines. Pupa like that of Eacles in type, ending in a large spine-like cremaster, and subterranean. There are three groups in this subfamily. The first and most generalized (Imbrasise) is represented by the three genera of which Vsta is the most generalized; while Cirina, and especially Imbrasia, with its tailed hind wings, is the most specialized. [Here also Gonimbrasia.] The second group is a more natural one, the Bunaese, all the three genera being closely allied, and then- larva? known. They are Thyella, [Acanthocampa] belina and Bunaea. The third is perhaps more modern, more specialized, some of the forms, as Nudaurelia, closely mimicking Antheraea of the Satumiidae, in the shape of the antennae, wings, and the presence of large ocelli similar to those of the silk-spinning family, though the larvae are very different as well as the transformations. USTA Wallengren. [Usta Wallengren, Wien. Ent. Mon., VII (1863), p. 142.] [Type Usta wallengrenii (Felder).] Imago. — s. Head moderately wide in front; fairly prominent, scales long, erect, adding to the size of the head and prominence of the front. Antennae of the male unipectinate, well pectinated, nearly to the tip, the last five joints without pectinations; the joints numerous (50), very short, only one pair of (basal) pectinations present with no vestiges of distal pairs which are naked, scarcely ciliated. Palpi very stout, thick, and short, only reaching to the front; consisting of but a single joint, small and short. Thorax moderately stout. No tongue visible even after the removal of a palpus. Fore wings rather narrow and small, of nearly the same proportions as in Saturnia; costa straight at base, much curved toward the apex, wliich is much rounded, unusually so; outer edge not so long as the inner edge, straight, not incurved. Hind wings moderately wide; apex weU rounded, outer edge not very convex; abdomen not quite reaching the inner angle of the hind wings. Venation: Vein II, arises not far beyond the middle of the discal cell, and close to the origin of the stalk giving rise to II3 and II4. Vein II2 present. Vein III2, not forming an inde- pendent and giving rise to the anterior discal vein; the two discal veins directed inward, forming a common line bent inward. Hind wings with the discal cell very long, the discal veins being situated far beyond the middle of the wing, the two together being sharply bent inward, the dis- tance between the origin of veins II and III being the same as between the origin of the latter and of the posterior discal vein. 30 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, Markings : Ground color white, spotted, or marbled with red ; a large round discal ocellus of nearly the same size in each wing. Hind wings nearly white. Fore legs with two large stout tibial spines at the end of the fore tibia, the outer spine larger than usual; the tibia is unusually short, compared with the femur. The odoriferous appendage is about three-fourths as long as the tibia itself, narrow, subacute, and hairy on the inside. This genus is SaturniaAike in shape and size of wings, as well as the discal ocelli, but the venation shows plainly enough family distinctions; it is remarkable for the very short bushy palpi, which are only one-jointed, and for the antennae which are not bipectinate, the discal pair of pectinations entirely wanting, there being no vestiges of them. Geographical distribution. — British East Africa, Tzavo. USTA ANGULATA Rothschild. TJsta angulata Rothschild, Novitates Zool., II, p. 50, PI. X, fig. 5, 1895. Imago. — One 6* . Body and wings a ground color of white scales, spotted and marbled with scattered bright roseate pink red scales. Head in front and fore legs tawny; breast roseate. There are on the body two roseate lines, one on each side of the thorax, also a transverse roseate line across the basal abdominal segment; the abdomen being white marbled with pink red. The scales arc very fine and close. Fore wings wliite, with scattered pink scales. An extradiscal very deeply and sharply zigzag line. Discal ocellus a black ring inclosing a bright ochre-yellow area, with a reddish center; the same on the hind wings, which are crossed by a wavy extradiscal line. The mark- ings are the same beneath. Expanse of fore wings, 9 92 mm. Length of fore wing, 9 45 mm. Breadth of fore wing, 9 23 mm. Length of hind wing, 9 32 mm. Breadth of hind wing, 9 24 mm. Described from a much rubbed imperfect specimen, received from the British Museum, with the examples in which my specimen was compared and named. Geographical distribution. — "British West Africa (Gregory coll.), Tzavo (from British Mu- seum); Mombasa, East Africa, lat. 4° 5' (Rothschild)." Rothschild states that this species differs from U. waUengreni: First, "the transverse angu- lated submarginal band in U. waUengreni (Feld.), is convex, follows the outline of the wings, and its angulations are the same size throughout, while in angulata the band is quite zigzag and the lower angulations are quite three times the size of the upper; secondly, the ocelli are much larger, and the fulvous center is reduced to a narrow ring. " "Expanse 3£ inches = 88 mm." ■ USTA WALLENGRENI (Felder). Salurnia waUengreni Felder, Wien. Ent. Mon., Ill, p. 323, Taf. VI, fig. 2, 1859. TJsta waUengreni Wallengren, Wien. Ent. Mon., VII, p. 142, 1863. Saturnia terpsichorina Westwood, Oatcs, Matabele Land, p. 357, 1881. Saturnia waUengreni Butler, Trans. Ent. Soc. London, 1889, p. 391. Heniocha terpsichorina Kirby, Syn. Cat. Lep. Hot., I, p. 771, 1892. TJsta waUengreni Rothschild, Nov. Zool., II, p. 50, 1895. [Geographical distribution, Caffraria]. — Fifty miles inland [from] Mombasa (Tring Museum). USTA (?) TERPSICHORE Maassen. Plate XXXI, fig. 13. Saturnia (?) terpsichore Maassen and Weymer, Beitrago Schmett., fig., $ , 113, 9, 114, 1885. Heniocha terpsichore Kirby, Syn. Cat. Lep. Het., I, p. 771, 1892. Heniocha terpsichore Rothschild, Nov. Zool., II, p. 49, 1895. Imago.— One 9 • Head a little darker, more fulvous than the thorax, which is of a soft pale leonine hue or fawn color, with no pink scales. Antennae well pectinated. Palpi deep red, as also the fore legs, except the last tarsal joint which is fawn color. no. i. BOMBYGTNE MOTHS OF NORTH AMERICA— PACKARD. 31 Fore wings subacute at tip, apex not rounded; the ground color whitish ash, reddish on the costal edge, especially beneath; the basal line curved, bent outward on veins III and V. Extradiscal line sinuous, somewhat parallel with the outer edge and quite regularly scalloped, the seventh scallop or point acute; all nearly the same size. Beyond are three or four parallel faint wavy ashen or fawn colored lines on a whitish field; edge of the wing light fawn. Discal ocellus round, slightly oval, moderately large (7 by 5 mm.) consisting of a black ring inclosing a pale fawn area, the center of which is darker fawn, with white scales, densest on the inner side. Hind wings cream-white; base of wing clear white; a faint extradiscal line nearly touching the ocellus; it is sinuous and fades out toward the costa. Edge of the whig sparingly dusted with dark scales. Ocellus about a third smaller than that on the fore wing (6 by 5 mm.), not quite round, the dull ochreous ring is broader than in the ocellus of fore wing, and incloses a brown center, with white scales on the inner side (no red scales as figured by Maassen). Beneath as above, but fewer white scales in the ocelli. Abdomen; with reddish scales along the sides of the abdomen, forming a series of lateral red spots. Expanse of fore wing, 9 100 mm. Length of a fore wing, 9 50 mm. Breadth of a fore wing, 9 27 mm. Length of a hind wing, 9 35 mm. Breadth of a hind wing, 9 28 mm. The specimen in the Schaus collection of the American Museum of Natural History agrees well with Maassen and Weymer's figure, but there are no red scales in the ocelli. Geographical distribution. — Delagoa Bay (Schaus, Maassen). CIRINA Walker. Saturnia Westwood, Proc. Zool. Soc. London, 1849, p. 52. Bunaea Walker, Cat. Lep. Het. Br. Mus., VI, p. 1232, 1855. Cirina Walker, Cat. Lep. Het. Br. Mus., VI, p. 1382, 1855. Sculna Wallengren, OEfv. vet. Akad. Forh., XV, p. 139, 1858. Cirina Ktrby, Syn. Cat. Lep. Het., I, p. 763, 1892. Imago. — - mark, the stalk of the >- extending out along the extradiscal space. Beneath there are no Lines or other markings. Palpi not reaching the front, not very stout, depressed. Fore wings rather produced toward the apex and hind wings well rounded on the apex. Abdomen black-brown, with five white rings on the front edge of five segments; end of the abdomen snuff-yellow. The thorax is partly denuded [of hairs], but there are no traces of erect stiff flattened ones. Expanse of the fore wings, .), XVIII, p. 1, 1872. Actias isabellx Maassen and Weymer, Beitrages Schemtt., Ill, figs. 40, 41; 1873. Tropaea isabellx Kirby, Syn. Cat. Lep. Het., 1, p. 765, 1892. Graellsia isabellx Grote, Die Saturniiden, Mitth. Roemer Museum, VI, p. 26, 1896. LARVA. Girard, Metamorphoses Ins., p. 263, figs. 211, 212. Milliere, Icon, et Descr. Chenilles et Lep., Annales Soc. Linn. Lyon (Nouv. S6r), XVIII, p. 1, 1872. [Chapman, Entom. Record, 1902, p. 126.] Moth. — One a* , one 9 . (Others seen.) Body brown madder-red, abdomen striped or banded with pale ochreous fawn. Wings delicate pea-green ; veins widened by madder. Discal ocelli with an outer black ring, and on the inside of the discal veins a wide pale steel-blue semi- circle, within red-pink and outside of the vein a broad yellow semicircle. No intra and extra discal lines, but three submarginal brown lines, of which the inner is about one-half as wide as the middle one. Edge of wings all around madder red-brown. Beneath exactly as above, including the ocelli, though the blue is less distinct. Expanse of fore wings, c? 80; 9 82 mm. Length of fore wing, o* 44; 9 45 mm.; length of hind wings [I noted in the U. S. National Museum that yellow-green specimens were males, light bluish green females. Dr. W. T. M. Forbes assures me that this is a regular sexual difference.] 190 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, The edge of the hind wings is yellow, but on the inner angle the fringe is whitish. Ocellus not so long as wide (4£ by 5 mm.), with no traces of red scales either inside or outside of the discal veins. The under side of both pairs of wings whitish, and the ocelli of both pairs of wings are much smaller and paler than in the winter form, those of the hind wings being no larger than those of the fore wings. Length of fore wings, 50 mm. Breadth of fore wings, 31 mm. Length of hind wing, 70 mm. Breadth of hind wing, 33 mm. Length of tail, 28 mm. Breadth of tail, 7 mm. In neither of the two forms are there any traces of an extradiscal line. Climatic varieties or races. — While T. luna from Maine to Texas and Florida and through- out the United States north of Mexico offers so far as yet known no local or geographic races, the case appears to be different in Mexico and Central America. There, on the southern outskirts of its range, it seems to have been affected by climatic agencies, and to have varied locally in different directions and to have been differentiated into perhaps three local races or varieties, i. e., T. luna var. 1, dictynna; 2, azteca; and 3 truncatipennis. (1) Tropaea dictynna Walker, Cat. Lep. Het. Br. Mus., VI, p. 1264, n. 8 (1855). Walker's description gives no locality for his specimen, which was contained in Mr. Saun- ders's collection. He remarks: "This species much resembles T. luna, but may be distinguished by the band on the wings, by the not empurpled exterior border, by the fore wings, which have a less oblique and more straight exterior border, and by the hind wings, which have shorter tails." A 9 in the Oxford Museum, which agrees with Walker's description, was examined by me, thanks to the courtesy of the attendant in charge. In this specimen the fore wings are broad and obtuse at the apex; there is a pale greenish-brown distinct extradiscal line, situated a little more than half way from the base to the aj)ex of the fore wings, being nearer the outer edge of the wing than to the ocellus, and the line reappears on the hind wing.1 The ocelli are quite as in T. luna and those of the fore wings are nearly separated from then- stalk. The edge of the wings is ochreous yellow. The tails are longer than in T. luna (Walker says the tails are shorter). The hind wings are a little scalloped. The impression made upon me was that this form is not distinct from T. luna. I also examined with more care and deliberation the 3 , labelled T. dictynna in the British Museum, from Orizaba, Mexico, "at or near the volcano," with the following results: It differs from four a* and five 9 of T. luna, in the same drawer, in the distinct extradiscal brown band, which is wide and crosses the fore wings, not quite reaching either the costa or the hind edge. The shape of the wings and the length of the tails as in T. luna. Outer edge of both wings yellow and pale brown, as in T. luna. In the hind wings the outer edge is not so much scalloped as in most of the examples of T. luna. The wings are ochreous green, rather more so than in the examples of T. luna, and the squamation is rougher. Ocellus of the fore wing as in T. luna, but there is less white next to the vitreous center, but more red, otherwise the shape and coloration of the ocellus is the same. In the ocellus of the hind wings the yellow ring is deeper ochreous than in some T. luna, but of the same hue as in some others; the red ring is distinct. The ocelli in fact show no varietal differences. The moth is not quite so large as the largest T. luna. At the time of examination I concluded that T. dictynna is hardly a climatic variety of luna. That dictynna is only a banded form of luna, and liable to occur anywhere, is probable from the fact that the late Mr. S. L. Elliot has bred it in Brooklyn on the walnut, having raised 1 Mr. W. L. W. Field informs me that in examples bred by him at Milton, Mass., the bands appear after emergence from the coeoon as soon as the wings expand, but that after about one hour they disappear. ) no. i. BOMBYCINE MOTHS OF NORTH AMERICA— PACKARD. 191 from 400 to 500 of this species. He once remarked to me that the tail in the banded individ- uals is on the whole inclined to be shorter than in the unstriped individuals. On the other hand, he said, the pale thin green ones bred on leaves of the sweet gum (Liquidambar) have very long tails, no bands, and the fore wings edged with reddish-pink. Are there food varieties ? Mr. Field informs me that in examples bred by him at Milton, Mass., bands appear after emergence from the cocoon as soon as the wings expand, but that after about an hour they disappear. [The following was evidently written by Dr. Packard at a different time:] Mr. A. G. Butler, Annals and Mag. Nat. Hist., fifth series, IV, p. 356, 1879, remarks as follows concerning this form: "Linnaeus evidently confounded T. dictynna with T. luna; for although in the tenth edition of his 'Systema,' he quoted Catesby's figure of the North American insect and Petiver's rep- resentations of the same species as illustrations to his brief diagnosis, yet for the fuller descrip- tion given in the 'Museum of Ulrica,' he quoted Clerck's 'Icones' first of all; and the figure in the latter certainly represents the Mexican species, inasmuch as the wings are crossed by a well-defined discal stripe. It is probable that both descriptions are taken from the northern type." It would seem, since dictynna is scarcely a local race of luna, much less a "Mexican species," that Linne did not confuse the two forms, for individuals with wings crossed by a well-defined discal stripe, are Hable to occur anywhere, north or south, or in Central America.' (2) Tropaea azteca Packaed, Guide to Study of Insects, p. 298 (1869). [The original description is as follows :] "In the museum of the Peabody Academy is a closely allied and undesciibed species from the west coast of Guatemala, which we would call Actias azteca. It differs from A. luna on its much smaller size, expanding only 3i inches, and in the shorter fore wings, the apex being much rounded and with shorter veins, while the 'tails' on the hind wings are only half so long as those of A. luna. It also differs in having the origin of the first subcostal venule much nearer the discal spot than in A. luna, being very near that of the second subcostal venule. It is whitish green, with markings not essentially differing from those of A. luna." Originally described as a new and distinct species, I have again examined the single 9 type specimen, which has been discovered in the collection of the Cambridge Museum, pre- sented to that museum by the Peabody Academy of Science, Salem, Mass. It was described as quoted above as differing from A. luna in its much smaller size, and in the shorter fore wings, the apex being much rounded, "while the 'tails' on the hind wings are oidy half as long as those of A. luna." It is whitish green, with markings not essentially differing from those of A. luna. What we then said regarding the supposed difference in the venation does not now seem of much weight, as there do not seem to be any perceptible differ- ences. A reexamination of the type has resulted in the following observations : The type of azteca is similar in shape of wings and in the discal spots with their stalk to the summer form of an ordinary T. luna from New York, but it has decidedly shorter tails than any specimen I have yet seen, while the apex of the fore wings is much more rounded. There are no traces of a submarginal line, as seen in var. dictynna. The discal ocellus of the fore wings is small and narrow, in the center is a narrow lanceolate oval clear space, edged within with a pink semi- circle, succeeded by a broader black crescentiform semicircle, containing a linear strip of pale blue scales. It measures 4 by 3 mm. The stalk is well developed. The blue fine in the discal ocellus of the hind wings is not distinct; the spot measures 4£ by 4$ mm. Expanse of the fore wings, 9 90 mm. Length of a single fore wing, $ 45 mm. Breadth of a single fore wing, 9 26 mm. i [Dr. W. T. M. Forbes informs me that all the specimens of T. luna from southern Florida in the American Museum of Natural History are of the dictynna type, and on the average quite different from the northern form. They have the banded wings and shorter tails, but differ from Walker's account of dictynna in having more crimson color than the ordinary luna. This last character may be seasonal.) 192 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, Length of a hind wing, 9 52 mm., including the tail, which is 21 mm. long and 5 mm. wide in the middle. Breadth of hind wing, 9 24 mm. Geographical distribution. — This form was collected at Polvon, occidental department, West Nicaragua, near the Bay of Realejo, but inland between Corinto and Leon, near Cor- cuera, by John A. McNiel. Whether this is even a local race peculiar to Nicaragua or not remains to be seen, when more specimens are discovered; the tails vary greatly hi length, though it is noteworthy that both truncatipennis and azteca occur hi Central America, forms in which there is so much diverg- ence as regards the length of tail. (3) Tropaea truncatipennis Sonthonnax [Lep. Soie, II (1903), p. 19, PI. VII, Jalapa]; PI. L, fig. 2. One 3 , one 9 . This form differs in the smaller ocelli of both pairs of wings ; the ocelli of the fore wings are so small that they do not reach vein IV. Beneath, the ocelli are notice- ably smaller than in luna. The shape of the wings is also different from luna of Texas north- ward, the fore wings being a third smaller and rounder, the hind wings being less rounded and more elliptical. This is the chief and only tangible difference between the two forms. The stalk, its size, shape, and coloration is just as in T. luna. The general hue is the usual delicate pale pea-green shade, as seen hi most of the northern specimens. The outer edge of the wings of both pairs is deep reddish pink. The hind wings are less scalloped than hi the normal T. luna, and the fore wings tend to be slightly more falcate. There is, as in most of the exam- ples of T. luna, no trace of an extradiscal band or line or any other markings. It is of the same size, the same expanse of wings, and the legs and body as just as in T. luna. Truncatipennis: Expanse of fore wings, 3 123 mm.; 9 126 mm. Length of fore wings, 3 60 mm.; 9 63 mm. Length of hind whig, including tail, 86 mm. Length from discal vem to tip of tail, 3 9 62-64 mm. Ocellus of fore, wing, 4 by 4.3 mm. Ocellus of hind whig, 4.3 by 5.5 mm. Luna: Expanse of fore wings, 3 125 mm. Length of fore wings, 3 60 mm. Length of hind whig, including tail, 3 85 mm. Length from discal vein to tip of tail, 3 64 mm. Ocellus of fore wing, 3 5 to 5.2 by 4.5. Ocellus of hind wing, 3 5.6 by 5.1. If these characters hold good hi a larger number of specimens, truncatipennis may safely be regarded as a climatic variety of T. luna. [An apparently more recent note on T. truncatipennis, based on a female in the Godman- Salvin collection (British Museum), from Jalapa, indicates that this is probably a valid species.] Fore wings falcate, outer edge curved in, but no more so than in some 9 luna, which varies much, some being falcate and others with outer edge not excavated at all. Hind wings pointed at apex and outer edge not full, but straight and not scalloped. Outer edge of both wings broadly margined with reddish brown (magenta?), but two of luna as much so. Tails a little longer than hi luna; considerably longer than breadth of the hind wing. Third median vein arises nearer base of wing than in luna. Venation of fore wing as in luna. Ocelli of fore wing as in some luna, well pronounced, red, yellow, blue semicircle and black. Ocellus of hind wing not so large as in some luna, but well marked; black is heavy, as is red, the yellow outer portion is pale green, less ochreous than in Mexican (Orizaba) specimen. Outer edge and apex of hind wing is less scalloped and more acute than in any luna. All four ocelli beneath are larger than hi Orizaba luna and United States luna. It seems quite distinct from the Mexican no. i. BOMBYCINE MOTHS OF NORTH AMERICA— PACKARD. 193 c? dictynna. Sir George Hampson thinks it is a good species; as the larva differs from luna (Schaus), it may be regarded as a distinct species.1 A new climatic race (?) hred in Europe. — Tropaea luna, var. bollii (Wagner); Saturnia bolli Wagner, Tijdschr. voor Entom., 1875-76, XIX, p. xcvii-xcviii; Isis, etc. The <$ specimen in the Cambridge Museum reared in Switzerland differs from the Texan normal luna S in the antennae being very narrow, but little more than half as wide as usual; the submarginal brownish line on the fore wings is distinct; ocellus small, its outer edge indis- tinct; that of hind whig is much smaller than in natives of Texas, about one-fourth less in diameter, and the pink line is wanting in the ocelli of the hinder wings. The tail is rather short, and the moth is a little smaller than other Texan luna. The edge of both wings is yellowish as in the summer form of T . luna. [Other varieties of T. luna have been described as follows: (1) Variety rossi (Ross). Male pure white, expanding 3§ inches; female white with a light yellowish tinge. Near Toronto, Canada. (Classif. Cat. Lep. Canada, 1872.) (2) Variety rubromarginata W. T. Davis. A spring form hi New York State, having the costa red or reddish, and the outer margin of the wings "of a reddish or scarlet hue." (Psyche, June, 1912, p. 91. )]2 Life history of Tropica luna (Linne). The eggs were received from Mr. James Angus. Egg. — Oval-cylindrical, somewhat flattened. The shell is thick and tough, dark brown externally, but in places the brown is worn off, leaving a dull, sordid chalky whitish surface; the inside of the shell slightly bluish gray. The surface of the shell is seen under a Tolles triplet to be rough and finely granulated, and under a half- inch objective, the surface is seen to be closely granulated, the pits between the granulations being often confluent; rarely the raised bosses appear to be polygonal. Length 2.1 mm., breadth 1.8 mm. Eggs laid at Providence, June 14; hatched June 22-25. Larva. — Stage I. Length 6-8 mm. Some were observed hatching out between 11 and 1 o'clock p. m., June 15. Before entirely breaking out of the eggshell the tubercles on the anterior segments become erect, and the hairs radiate from them, but behind along the third thoracic and abdominal segments the tubercles were seen to be soft and flattened or appressed to the body and adhering in flaccid bundles. In S. cecropia, on the other hand, all the tubercles and bristles are flabby for perhaps hah an hour after the creature frees itself from the egg. One was seen to emerge at 1.15 p. m., and by 1.25 p. m. all the tubercles had become filled out and erect, with stiff, radiating bristles.3 On hatching, the body is entirely green, except the bands on the head. Some larva? on hatching are (a) entirely yellowish green, while the dorsal hairs are darkish, and the head is twice banded. Others (b) have a very broad blackish lateral band, inclosing one lateral row of greenish tubercles, the band ending on the eighth abdominal segment, and nearly meeting above. The prothoracic segment is dark on the hinder edge, and the second and third thoracic and first abdominal segments are entirely dark above. The following description is drawn up from individuals which had been hatched for about a week (May 24-26), and were near the end of this stage. The body was larger, fuller, and less tapering posteriorly than at first. The head is small, about half as wide as the body, rounded, and at rest can be retracted within the prothoracic segment. There is a transverse dark i [The U. S. National Museum has eight specimens of T . truncatipennis; the localities are Jalapa, Guadalajara, and Orizaba. They all have the outer margins of the wings narrowly red. The costo-apical angle of the secondaries seems normally much more acute than in luna, and from here to the base of the tail the margin is straight, whereas it is distinctly convex and wavy in luna. None shows any traces of dictynna bands. The sexes seem not to differ in color.] 2 [Dr. W. T. M. Forbes suggests that the account of rossi could apply to newly emerged examples of typical luna, with the descriptions of the sexes reversed. Excellent specimens of rubromarginata are in the National Museum, and also a more extreme ab. rubrosuffusa, of which a male was taken at Washington, D. C. (Knab). This is like rubromarginata, but has the red of outer wing margins, especially of the hind wings, broader and suffused, on hind wings at base of tails fully 5 mm. broad, and this is bordered within by a pallid, almost bluish suffusion. The ocelli on the hind wings are unusually large.] 8 It is evident that before and at the point of hatching the setse or bristles are filled with blood, which distends them. While thus distended, the fluid may ooze out of the ends, and thus they may be called glandular hairs. In those which are full and bulbous at the end, the fluid may be retained through stage I, and in rare cases through the second or even the third stage. 83570°— 14 13 194 MEMOIRS NATIONAL ACADEMY OF SCIENCES. Vol. XII, brown band in front just below the vertex, making two scallops, and ending on the sides; on each side (below) of the front edge of the clypeus is a dark spot around the base of the antennae, which sometimes sends a short line inward, as in Mr. Bridgham's figure. The body is thick, full, cylindrical, each segment, except the prothoracic and last two abdominal ones, with six thick, smooth conical tubercles, those on the sides above the spiracles smaller than those below, and about one-half the size of the dorsal ones, and bearing fewer bristles than the others. Prothoracic segment with only four tubercles, the two dorsal ones low, flattened, and small, with about 14 radiating bristles. The lateral tubercles are like those of the other segments; the rest of the dorsal tubercles are large, full, nearly touching at their base, and bearing about 8 to 10 bristles, which are one-half to one-third longer than the tubercles themselves; they radiate and are dark purplish, pale at base, those on the back darker than those arising from the lateral tubercles. The second and third thoracic dorsal tubercles are slightly larger than the abdominal ones. Each of the dorsal abdominal tubercles bears about six bristles. The body is delicate pea-green, nearly like the under side of the Carya leaf on which they feed. The tubercles, especially the dorsal ones, are tinged with faint straw or lemon yellow, while the lateral supra- spiracular tubercles are greenish, scarcely tinged with yellow. The bristles are longer in proportion to the tubercle than in the larva of C. promethea; most of them are three times and some four to five tunes as long as the tubercle. The bristles are sparingly and minutely barbed, tapering acutely, but they are clear, and per- haps glandular. The median dorsal tubercle on the eighth uromere shows traces of its double origin, but they are not so marked as in C. promethea and T. polyphemus, but more so than in Platysamia cecropia. It is much broader than long at base, and on the tip bears five setae on each side. The ninth uromere bears four tubercles of equal size, which are large and well developed, the lateral ones scarcely smaller than the dorsal ones. The suranal plate is broad and short, more so than in T. polyphemus, not tubercled, but bearing two tufts of bristles which are but a little shorter than those arising from the lateral tubercles of the rest of the body. The anal legs are large and squarish, as in the group generally; all the legs, both thoracic and abdominal, are pale green. The abdominal legs bear each 20 crochets. The three tenant hairs of the thoracic feet are rather longer than usual. The spiracles arc slightly chitinous, not colored. The shape of the double dorsal tubercle on the eighth abdominal segment is shown at figuro 18, d'; sd, the subdorsal ones; a, a seta much enlarged, which, unlike T. polyphemus, is finely and minutely barbed; a', a", ends of other setae. Stage II: Molted May 26, in the daytime. Length at first 9 mm., afterwards 10 mm. In one larva all the tubercles are of the same yellowish hue; in the other, those of the second and third thoracic segments are brownish at the tip, thus greatly contrasting with the others. In another larva the median dorsal tubercle on the eighth uromere is also colored in the same way. The head in one is all green, not yet banded with brown; but in another the head is partly banded, i. e., in place of the two-scalloped band are two separate short scallops. The tubercles are now higher than before, and rough with slender conical warts which give origin to the setae. The prothoracic tubercles are now longer than before, and all four are deep amber-yellow at the end, the setae being black; two out of the five spines of the second and third thoracic segments are dark brown at and near the ends and give rise to black bristles, rendering Fig. 18. — Tropsea luna; larva, first stage. no. i. BOMBYCINE MOTHS OF NORTH AMERICA— PACKARD. 195 them very conspicuous ; they are a little larger and higher than those on the abdomen, and bear about twice as many bristles; eight in all, all of which are black, while on the yellow tipped tubercles of the abdominal segments there are about five bristles, one of them minute; two of the five are black, the others pale. The two lateral rows of tubercles are, as before, with pale bristles. The median dorsal tubercle on the eighth uromere is not quite so dark as those on the second and third thoracic segments, and some of the latter are scarcely darker than the other abdominal ones. The spiracles are of the same pale color as before. The suranal plate still bears the two terminal tubercles, as before. The thoracic legs are now darker than before. In this stage the larvae sometimes assume a sphinx-like attitude. Stage III: Molted June 1. (I am not sure that it was the same larva; one molted May 31. Described three days after molting.) Length 13 and finally 15 mm. The head is either banded as before, or all green, only the ocelli being black. The body is now thick, though differing very slightly from the preceding stage. The four prothoracic, the two dorsal second and third thoracic tubercles, and the single median dorsal tubercle on the eighth uromere are either deep crimson red at the end, or much paler, and in the largest one yellowish, the tips of these tubercles varying a good deal in color; these tubercles are now nearly twice as long and thick as those on abdominal segments 1 to 7 and 9. The tubercles of the two lateral rows are of the same size as before; those of the upper (supraspiracular) row are stUl green and small; those below, situated on the lateral ridge, are salmon-colored and provided with black setae, like those arising from the dorsal tubercles; near, and on the base of and between the tubercles, are white, delicate clavate hairs (glandular?) which are not observable in the preceding stage; they are mostly confined to the abdominal, few, only one or two, on the thoracic region. The thoracic legs are dark brown, pale at the tip; the abdominal legs except the anal pair, are green, with a transverse lilac line near the ends; beyond yellowish, while the plantae are tinged with lilac. There is as yet no lilac tinge on the edge of the suranal plate- Stage TV: (Belonging to a later different brood; described July 24.) Length 23 mm. The head now pea-green, not banded in front, nearly as wide as the body; well rounded, and of the same shape as in T. polyphemus; it is of a deeper pea-green than the body, which is in general, especially on the dorsal side, paler than in T. polyphemus. The labrum and jaws are pale. There is a chestnut-colored ocellar patch. The segments are now quite convex, swollen under the base of the tubercles, the second and third thoracic segments being fuller and more angular than the uromeres; they are a little more so than in Telea polyphemus. The four dorsal tubercles of the second and third thoracic segments (two each) are larger than the abdominal ones, and are tipped with dark carmine at the end, and each, besides one or two short setae, bears a long black slender hah, about as long as the body is thick; the corre- sponding hairs on the abdominal tubercles being about one- third as long. There are four well- developed prothoracic tubercles, the dorsal ones larger, more rounded, and prominent than in T. polyphemus, and also bearing besides three or four small, short pale hairs, and a black very long one. The prothoracic tubercles are deep rosy pink, not coral-red. The lateral ones on the same segment are nearly twice as large as those behind in the same series, and all on the body are rosy pink or "crushed strawberry" color. The lateral infraspiracular ridge along the abdominal segments is distinctly lemon-yellow. The spiracles are faint reddish green, quite inconspicuous. The thoracic legs are reddish. The middle abdominal legs are green above, below is a narrow distinct black stripe, the end yellow, while the planta is livid flesh color; the anal legs with an anterior oblique yellow band, and a black spot corresponding to the black stripe, with black hairs above, as on the middle legs. The suranal plate is faintly edged with yellow. The larva in this stage differs from T. polyphemus of the same age in the green head, the distinct lateral yellow stripe below the spiracles, which are green, and not readily seen. The six dorsal thoracic tubercles are distinctly larger and more prominent than the abdominal ones, and they each bear a single very long slender black hair, besides one or two short ones; this is a 196 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xii, good generic character, separating it at once from T. polyphemus, and the suranal plate is not edged with purple, but with faint yellow. When fully fed,1 its length is 65 mm. Maine, August 20. The head is green, of a different hue from the body, more like Paris green. The body is large, heavy, plump, and thick, much as in T. polyphenols, and the tubercles are pinkish red, or crushed strawberry. The suranal plate is edged with yellow in front, but the surface is coarsely granulated, and in color dull amber; there is a similar long narrow patch on the side of the anal legs, bordered above with black and straw-yellow. The spiracles are green with the edge of the linear opening ochreous. The yellow lateral line is obscure. The body is still provided with white hairs, not arising from tubercles. The body is pea-green, dorsally slightly tinged with ruddy. Recapitulation of the more salient ontogenetic features. A. Congenital features. 1. Setae tapering to a point, not bulbous, and finely barbed. Stage I. Most of them are three or four times as long as the tubercles. 2. Some larvae in stage I with a very broad lateral dark band along the side of the body, some without it ; no transverse stripes present, but the head in front is twice banded with dark brown. 3. The 2d and 3d dorsal thoracic tubercles differentiated in stage I, being slightly larger than the abdominal ones. 4. On the suranal plate are two rudimentary tubercles, each bearing a tuft of bristles. 5. The dorsal median tubercle on uromere 8 does not show such marked traces of its double origin as stage I of C. promethea, or T. polyphemus, but it is more duplex than in S. cecropia. B. Evolution of later Adaptational Characters. 1. Dorsal tubercles in stage II higher than before. 2. The lateral dark band disappears in stage II. 3. In stage III the dorsal thoracic tubercles become nearly twice as long and thick as the abdominal ones. 4. The head is not banded in stage IV. 5. The tubercles brightest (pink or dark carmine) and most conspicuous in the last stage. 6. A distinct infraspiracular yellow line in stage IV, and the suranal plate and anal legs lined with yellow, and the surface of the suranal plate and sides of the anal legs amber. [The records of the United States Department of Agriculture indicate the occurrence of T. luna in the following localities : New Hampshire (Derry, Center Sandwich) ; Vermont (Nor- wich) ; New York (Brooklyn) ; New Jersey (Hackensack, Allendale) ; Pennsylvania (Spring Creek, Lancaster, Tamaqua) ; OMo (Franklin) ; Maryland (Pearson) ; Virginia (Carlin Springs) ; North Carolina (High Point, Wilkesboro, Raleigh) ; Kentucky (Louisville) ; Illinois (Douglas) ; Arkansas (Orlando) ; Indiana (Rockville) ; Tennessee (Henry) ; Louisiana (New Orleans, Wilsons Point) ; Mississippi (Craig) ; Florida (Astor, Madison, Kissiminee) . I picked up a fragmentary specimen from the seabeach at Woods Hole, Mass. The following are from the records of the Bureau of Entomology: V. T. Chambers counted 22 Tachina eggs on a single T. luna larva; this larva produced a moth, notwithstanding. (C. V. Riley.) A. H. Mundt, of Fairbury, 111., sent eggs of T. luna which were infested with what was supposed to be a species of Pteromalus. (T. Pergande).] [Prof. M. II. Swenk reports that T. luna "is common in eastern Nebraska wherever walnuts occur. We have specimens from Lincoln, Nebraska City, Weeping Water, Beatrice, etc." Gitt July, 1912).] "Not uncommon from here [Ottawa] westward throughout Ontario, and up to Winnipeg in Manitoba. I have seen two specimens in Manitoba, but doubt if it extends farther than the wooded country on the east of that Province. There are no maples, butternuts, hickories, or walnuts native in Manitoba." (J. Fletcher, litt 1900.) 1 Dyar states that there are hut four stages. no. i. BOMBYCINE MOTHS OF NORTH AMERICA— PACKARD. 197 ANTHER^A Hiibner. [Antherxa Hiibner, Verz. bek. S^hmett. (1S22?), p. 152. Type, according to Kirby, A. nylitta Drury.] [Rothschild, Nov. Zool., II (1895), pp. 43-44, arranges the species thus:] 1. A. assamensis (Helf.) 2. A. Tielferi Moore. [Darjiling.] 3. A. perrotteti (Guer.). [Pondicherry.] 4. A. papliia (Linn.). [Java, Ainboina.] ' 5. A. andamana Moore. [Andaman Islands.] 6. A. mylitta (Drury). [India.] ab. cingalesa Moore. 7. A. semperi Feld. [Luzon, P. I.] 8. A. pernyi (Guer.). [North China.] subsp. roylei Moore. [Darjiling.] ab. confuci Moore. [Shanghai.] ab. sJierviUei Moore. 9. A. larissa (Westw.). [Java.] 10. A.fritTiii Moore. [Darjiling.] ab.fraterna Moore. [Himalayas.] 11. A. billitonensis Moore. [Billiton Island.] 12. A. yama-mai (Guer.). [Japan.] [Rothschild states that no two specimens are alike, and treats the following variations or aberrations as synonyms: sergestus (Westw.), rnorosa Butl. Jutzina ~Butl.,f entoni Butl., calida Butl.] 13. A. sciron (Westw.). [Waigiou.] 14. A. pristina Walk. [Papua.] [Jordan (1910) described A. castanea from Assam. Many other species have been described by Moore (1892), Swinhoe (1892, 1893), Hampson (1892), and Lucas.] [Watson, Wild Silk Moths of the World (Manchester, 1912), has given colored figures of A. yama-mai (GueY.), A. compta Rothschild, A. mylitta (Drury), A. pernyi (Guer.), and A. anda- mana Moore. He also figures the cocoons of all of these except compta. A. andamana is a curi- ous plum-colored species, the wings with the ocelli relatively small, the anterior wings of the male very strongly falcate.] ANTHERXA YAMA-MAI Guerin. Plates XXVI, figs. 2-5; XLVII, figs. 4-6. [Bombyx yama-mai Guerin, Rev. Zool. (2) XIII (1861), p. 435; Pis. 11-13.] I am indebted to Prof. Sasaki for the eggs, which hatched at the end of March. Larva. — Stage I: Freshly hatched. Length 6 mm. Head large, chestnut red. Body thick, of the general saturnian shape of the larva of Telea, etc., at the same age. Body straw- yellow, with a narrow black and a lateral or subdorsal black stripe, the latter a little narrower than the dorsal stripe; this dorsal stripe stops at the median tubercle on the eighth abdominal segment, while behind this tubercle is a transverse black line or suture between the eighth and ninth segments. The setae or hairs are at first brown, very soon becoming black, but the tubercles at first are all yellow; afterwards they begin to turn black at the base, the ends remaining yellow; at the end of the stage they all become black. Thus, when from 24 to 36 hours old, the two dorsal tubercles on the third thoracic tubercle become black, as also the median one on the eighth abdominal segment. There is also a black triangular patch on the suranal plate, and one on the side of each anal leg. The tubercles of the second and third thoracic segments are very large and conical; the median tubercle of the eighth abdominal segment is double, high; the two dorsal ones on the ninth segment are high, and the supraspiracular tubercles are prominent. ' [The A. paphia of Kirby's Catalogue, from West Africa, is Nudaurelia dione (Fabr.).J 198 MEMOIRS NATIONAL ACADEMY OF SCIENCES. vol. xn, In the freshly hatched larva the black dorsal or heart line is heavier and more distinct than in the larva at the end of this stage. Cervical plate of the prothoracic segment gamboge-yellow, the front edge around each sub- median tubercle whitish. The hairs above are brown-black, while low down on the sides of the body, on the head and prothorax, they are whitish. The dorsal setae are rather stout, with slightly marked short spinules, while the setae of the lateral tubercles (supraspiracular and infraspiracular series) are slenderer, with longer delicate spinules, and longer than in Telea polyphemus, stage I, being about three times as long as the tubercle itself. Larva. — Fully grown. June 18, 1901. Length 75 mm. Of the size and general appear- ance of Telea polyphemus, but the segments are not so angular and distinct. Head large, rounded, nearly one-half as thick as the body, smooth, with pale hairs of uneven length in front upon and on each side of the clypeus ; green a little darker than the body, which is of a delicate pea-green, the hue of the underside of oak leaves. Prothoracic segment lunate, the cervical plate yellowish on the front edge, and behind greenish yellow. A few minute hairs on the front edge, but no warts, except a small 3-setiferous dark green flattened one low down on each side of the same segment. Second and third thoracic segments decidedly convex, the two dorsal tubercles, one on each side, fleshy, cmite distinct, giving rise to two or three long black hairs, and four short, rather sharp stout setae; the two tubercles are connected by a dark dusky shade. There are similar but smaller tubercles on the first and second abdominal segments with six straw-yellow hairs arising, two in a line across the segment, one dark one on each tubercle, and three or four spines. On abdominal segment 3-9 the tubercles are nearly obsolete, bearing only one or two hairs. While the dorsal and supra- spiracular lateral tubercles are green and of the same hue as the body, those of the lowest infra- spiracular row are turquoise blue; two on the side of the second and third thoracic segments, the lowest being situated just above the base of the legs. On the side of the second and third thoracic segments, just above each spiracle, is a conical metallic polished silvery smooth tubercle, each bearing a slender black hair ; the tubercle on the second segment is a little larger than the one behind it. The skin is dorsally and on the sides covered with scattered minute clavate whitish setae, as usual in the larvae of this family. On the eighth abdominal segment is a small but distinct tubercle, with two or three spines on each side, showing its double origin. The yellow hairs are about one-third as long as the body is thick. Spiracles pale yellowish testaceous. A lateral whitish yellow line just above the spira- cles and slightly edged above with livid or liver brown; on the eighth abdominal segment the liver-colored brown predominates, and becomes the very distinct livid brown of the edge of the suranal plate and sides of the anal legs. The thoracic legs are pale, the abdominal ones all green except the anal in part; the abdominal legs bear on the sides black and white hairs. Cocoon. — Regularly oval, no stalk, the surface like that of Bombyx mori; a large opening in front; color a rich greenish yellow. Length 43 mm.; thickness 22 mm. ANTHER4EA PERNYI Guerin. Plate XXVII. [Saturnia pernyi Guerin, Rev. Zool., 1855, pp. 6, 297; PI. 6, fig. 1.] The tip of the male antenna? of A. pernyi and yama-mai end in a simple point, with rudi- mentary pectinations, more so than in T. jwlyphemus. The fore wings of 9 are less falcate than in A. yama-mai. The dislocated basal band is redder than in A. yama-mai, and the extradiscal band is distinctly red, while that of A. yama-mai is dusky brown, and the band beyond is distinctly white, while the same portion of the band in A. yama-mai is narrower and pinkish. The ocellus on the fore wing of